I'm going to say either no, or that their body fur covers them. After all, the only reason human males have nipples is because our biology couldn't be bothered to get rid of them after the embryo stage. It could be mobians are evolved that males don't have nipples.
I see. So, how many nipples the females have?
Because different animal have different pair too.
For examples foxes have eight nipples arranged in four pairs, but hedgehogs have five pairs of nipples, totaling ten nipples,
I wouldn't be surprised if Sega forgot about him. I mean, with such puzzle games out there like Bomberman, Dig-Dug or Bubble Bobble, you'd think Sega would have Pengo compete against those games.
For me the problem because they changed the core gameplay.
Pepenga Pengo differs significantly from the original Pengo.
I know they try to innovate but sometimes simplicity is the best
(take Tetris, Frogger, and Pac-Man for example with Tetris 99, Frogger in Toyland, and Pac-Man 256)



and I do like Pengo's original gameplay.
And I guess that's the reason too why Switch version's gameplay is also back to its root.

Does sega still think Shadow has no friends? Or doesn't regard Rouge as one?
I think Shadow is a taundere.
While may be he said Rouge or Omega aren't his friends, but he still gonna helped them if he's needed.
His words and actions sometimes didn't match.
